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Position: Research Director
Expertise: Biologist Oceanographer
Department: Biological Oceanography
Premises: athens
Epaminondas D. Christou is a Research Director in the Institute of Oceanography of HCMR. He received B. Sc. (1981) from the University of Thessaloniki, M. Sc. (1984) and Ph. D. (1991) from the University of Athens. Since 1988, he has held a position as a researcher in HCMR and since 2003 as a Research Director. He has at shorter periods worked at: Station marine d'Endoume, Marseille, Kristineberg Marine Research Station, Rosential School of Marine and Atmospheric Science (RSMAS), University of Miami. He has a research interest in zooplankton (especially copepods and jellyfish) ecology, physiology and production having worked in several locations (Mediterranean, Skagerrak, Swedish fjord) on fluxes associated to zooplankton. He has acted as coordinator in national and international projects of HCMR. He has been involved in scientific and organizing committees as well as in organizing international scientific workshops (committee member or coordinator). He has published over 70 papers, 50 of which in peer-reviewed Journals, and has supervised several PHD theses.

Selected publications:

Siokou I., S. Zervoudaki, D. Velaoras, A. Theocharis, E. D. Christou, M. Protopapa, M. Pantazi. (2019). Mesozooplankton vertical patterns along an east-west transect in the oligotrophic Mediterranean sea during early summer, Deep Sea Research Part II: Topical Studies in Oceanography, 164, 170-189,
Kasapidis P., I. Siokou, M. Khelifi-Touhami, M.G. Mazzocchi, M. Matthaiaki, E. Christou, M.L. Fernandez de Puelles, A. Gubanova, I. Di Capua, S. Batziakas, C. Frangoulis. (2018). Revising the taxonomic status and distribution of the Paracalanus parvus species complex (Copepoda, Calanoida) in the Mediterranean and Black Seas through an integrated analysis of morphology and molecular taxonomy, Journal of Plankton Research, 40(5), 595–605,
Christou, E.D., Zervoudaki, S., De Puelles, M.L.F., Protopapa, M., Varkitzi, I., Pitta, P., Tsagaraki, T.M., Herut, B. (2017). Response of the calanoid copepod clausocalanus furcatus, to atmospheric deposition events: Outcomes from a mesocosm study Frontiers in Marine Science, 4:35 doi: 10.3389/fmars.2017.00035.
Berraho, A., Abdelouahab, H., Charib, S., Essarraj, S., Larissi, J., Abdellaoui, B., Christou, E.D. (2016). Copepod community along the Mediterranean coast of Morocco (Southwestern Alboran Sea) during spring,. Mediterranean Marine Science, 17/3, 661-665, doi:
Siokou I., S. Zervoudaki, E. D. Christou,(2013): Mesozooplankton community distribution down to 1000 m along a gradient of oligotrophy in the Eastern Mediterranean Sea (Aegean Sea). Journal of Plankton Res 35(6), 1313–1330, doi:10.1093/plankt/fbt089.
Zervoudaki, S., C. Frangoulis, C. Svensen, E. D. Christou, E. Tragou, E. G. Arashkevich, T. N. Ratkova, Ι. Varkitzi, E. Krasakopoulou, K. Pagou, (2013): Vertical Carbon Flux of Biogenic Matter in a Coastal Area of the Aegean Sea: The Importance of Appendicularians. Estuaries and Coasts, DOI 10.1007/s12237-013-9723-z.
Isari, S., Somarakis, S., Christou, E., & Fragopoulu, N. (2011). Summer mesozooplankton assemblages in the north-eastern Aegean Sea: The influence of Black Sea water and an associated anticyclonic eddy. Journal of the Marine Biological Association of the United Kingdom, 91(1), 51-63. doi:10.1017/S0025315410000123.
Zervoudaki S., E.D. Christou, G. Assimakopoulou, H. Örek, A.C. Gucu, A. Giannakourou, P. Pitta, T. Terbiyik, N. Yucel, T. Moutsopoulos, K. Pagou, S. Psarra, E. Özsoy and E. Papathanassiou (2011). Copepod communities, production and grazing in the Turkish Straits System and the adjacent northern Aegean Sea during spring. Journal of Marine Systems 86 (3-4), 45-56.
Molinero J. C., V. Vukanic, D. Lucic, F. Ibanez, P. Nival, P. Licandro, A. Calbet,  E. D. Christou, N. Daly-Yahia, M. L. Fernandez de Puelles, M. G. Mazzocchi, I. Siokou-Frangou (2009). Mediterranean marine copepods: basin-scale trends of the calanoid Centropages typicus. Hydrobiologia, 617 (1), 41-53.
Siokou-Frangou, S. Zervoudaki, E. D. Christou, V. Zervakis, D. Georgopoulos (2009) Variability of mesozooplankton spatial distribution in the North Aegean Sea, as influenced by the Black Sea waters outflow. Journal for Marine Systems, 78( 4), 557-575.
Economou G., E.D. Christou, A. Giannakourou, E. Gerasopoulos, D. Georgopoulos, V. Kotoulas, D. Lyra, N. Tsakalis, M. Tzortziou, P. Vahamidis, E. Papathanassiou, A. Karamanos (2008). Eclipse effects on field crops and marine zooplankton: the 29 March 2006 total solar eclipse. Atmospheric Chemistry and Physics 8, 4665-4676.
Zervoudaki S., E. D. Christou, T. G. Nielsen, I. Siokou-Frangou, G. Assimakopoulou, A. Giannakourou, M. Maar, K. Pagou, E. Krasakopoulou, U. Christaki and M. Moraitou-Apostolopoulou (2007). The importance of small-sized copepods in a frontal area of the Aegean Sea. Journal of Plankton Research 29(4), 317–338.
Shiganova TA, Christou ED, Siokou-Frangou I (2007) First finding of alien species Beroe ovata Mayer 1912 in the Aegean Sea. Mediterranean Marine Science 8:5–14.
M G Mazzocchi, E D. Christou, I Di Capua , M L Fernandez de Puelles, S Fonda-Umani, J C Molinero, P Nival, I Siokou-Frangou (2007). Temporal variability of Centropages typicus in the Mediterranean Sea over seasonal-to-decadal scales. Progress in Oceanography, 72, 214–232.
Siokou-Frangou, I., Sarantakos, K. and Christou, E.D. (2006) First record of the scyphomedusa Rhopilema nomadica Galil, 1990 (Cnidaria: Scyphozoa: Rhizostomeae) in Greece. Aquatic invasions, 1, (3) pp 194-195
Zervoudaki S., T.G. Nielsen, E. D. Christou and I. Siokou-Frangou (2006). Zooplankton distribution and diversity in a frontal area of the Aegean Sea. Marine Biology Research, 2:3, 149 – 168.
Tönnesson K , Maar M, Vargas C, Møller EF, Satapoomin S, Zervoudaki S, Christou E, Giannakourou A, Sell A, Petersen JK, Nielsen TG, Tiselius P. (2005) Selective feeding and grazing impact of Oikopleura dioica on an autmn protist community. Marine Biology Research 1(5): 365-373.
Maar M, Nielsen TG, Gooding S, Tönnesson K, Tiselius P, Zervoudaki S, Christou C, Sell A and Richardson K (2004) Trophodynamic function of copepods, appendicularians and protozooplankton in the late summer zooplankton community in the Skagerrak. Marine Biology, 144: 917–933
Tiselius P, J Petersen, TG Nielsen, M Maar, E Moller, S Satapoomin, K Tönnesson, T Zervoudaki, E. Christou, A Giannakourou, A Sell, C Vargas (2003). Functional response of Oikopleura dioica to house clogging due to exposure to algae of different sizes. Mar. Biol 142: 253–261
Maar, M, T G Nielsen, K Richardson, U Christaki, O S Hansen, S Zervoudaki, E D Christou (2002). Spatial and temporal variability of food web structure during the spring bloom in the Skagerrak. Mar. Ecol. Prog. Ser 239: 11–29
Vargas CA, Tönnesson K, Sell A, Maar M, Møller EF, Zervoudaki T, Giannakourou A, Christou E, Satapoomin S, Petersen JK, Nielsen TG, Tiselius P (2002). Importance of copepods versus appendicularians in vertical carbon fluxes in a Swedish fjord. Mar. Ecol. Prog. Ser 241:125-138.
Siokou-Frangou,I., Bianchi,M., Christaki,U., Christou, E.D., Giannakourou,A., Gotsis, O., Ignatiades,L., Pagou,K., Pitta,P.,Psarra,S., Souvermezoglou,E., Van Wambeke,F, Zervakis,V. (2002). Carbon flow in the planktonic food web along a gradient of oligotrophy in the Aegean Sea (Mediterranean Sea). J. Mar. Sys. 33-34: 335-353
Shiganova,T., Mirzoyan, Z., Studenkina, E., Volovik, S., Siokou-Frangou, I., Zervoudaki, S., Christou, E.D., Skirta, A., Dumont, H. (2001) Population development of the invader ctenophore Mnemiopsis leidyi in the Black sea and in other seas of the Mediterranean basin. Mar.Biol. 139, 431-445
Fragopoulu N., Siokou-Frangou, I, Christou E.D., Mazzocchi M.G. (2001). Patterns of vertical distribution of Pseudocalanidae and Paracalanidae (Copepoda) in pelagic waters (0-300 m) of the Eastern Medirerranean Sea. Crustaceana 74: 49-68.
Siokou-Frangou,I., Gotsis-Skretas,O., Christou,E.D. & Pagou K. (1999) Plankton characteristics in the NW Levantine Sea and the adjacent areas. In: Malanotte-Rizzoli P. and Eremeev V.N. (eds) The Eastern Mediterranean as a laboratory basin for the assessment of contrasting ecosystems. Kluwer Academic publishers. 205-223
Christou, E.D. & Stergiou, K. (1998). Modelling and forecasting the fortnightly cladoceran abundance in the Saronikos Gulf (Aegean Sea). J. Plankton Res. 20: 1313-1320.
Christou, E.D. (1998). Interranual variability of copepods in a Mediterranean coastal area (Saronikos Gulf, Aegean Sea). J. Mar. Sys., 15: 523-532.
Ban S, Burns C, Castel J, Chaudron Y, Christou E, Escribano R, Umani SF, Gasparini S, Ruiz FG, Hoffmeyer M, Ianora A, Kang HK, Laabir M, Lacoste A, Miralto A, Ning X, Poulet S, Rodriguez V, Runge J, Shi J, Starr M, Uye S, Wang Y (1997). The paradox of diatom-copepod interactions. Mar. Ecol. Prog. Ser. 157:287-293.
Siokou-Frangou, I., Christou E.D., Fragopoulu N.,. Mazzocchi M.G.(1997). Mesozooplankton distribution from Sicily to Cyprus (Eeastern Mediterranean): II. Copepod assemblages. Oceanol. Acta, 20: 537-548.
Mazzocchi, M.G., Christou E.D., Fragopoulu N., Siokou-Frangou I (1997). Mesozooplankton distribution from Sicily to Cyprus (Eeastern Mediterranean): I. General apects. Oceanol. Acta, 20: 521-535.
Christaki, U., F. Van Wambeke, E.D. Christou, P. Conan & R. Gaudy (1996). Food web structure variability in the surface layer, at a fixed station influenced by the North Western Mediterranean Current. Hydrobiologia, 321: 145-153.
Christou, E.D. & M. Moraitou-Apostolopoulou (1995). Metabolism and feeding of mesozooplankton at the eastern Mediterranean (Hellenic coastal waters). Mar. Ecol. Prog. Ser. 126: 39-48.
Christou, E.D. & G.C. Verriopoulos (1993). Length, weight and condition factor of Acartia clausi (Copepoda) in the eastern Mediterranean. J. mar. biol. Ass. U.K. 73: 343-353.
Christou, E.D. & G.C. Verriopoulos (1993). Analysis of the biological cycle of Acartia clausi (Copepoda) in a meso oligotrophic coastal area of the eastern Mediterranean Sea using time-series analysis. Mar. Biol. 115: 643-651.

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